(1809-1882)

Charles Darwin was born in Shrewsbury, England, on February 2, 1809.  His father was Robert Waring Darwin, a physician and son of the famous Erasmus Darwin, also a physician, as well as a respected writer and naturalist.  His mother was Susannah Wedgwood Darwin.  She died when Charles was eight. 

Charles was educated in the local school, taught by Dr. Samuel Butler.  In 1825, he went to Edinburgh to start studying medicine, but he soon realized that he did not have the stomach for it!  So he switched to Cambridge, ostensibly to become a clergyman.  He was actually more interested in entomology -- especially beetles -- and in hunting.  He graduated from Christ’s College in 1831.

It is said that even when he was a young man, he had a patient and open mind, spending many hours collecting specimens of one sort or another and pondering over new ideas.  The idea of evolution was very much in the air in those times:  It was increasingly clear to naturalists that species change and have been changing for many millennia.  The question was, how did this happen?

One of his mentors, John Henslow, encouraged him to apply for the (unpaid!) position of naturalist on a surveying expedition on the now-famous vessel, the Beagle, under the command of Capt. Robert Fitz-Roy.  Charles left England for the first time in his life on December 27, 1831.  He wouldn’t return until October 2, 1836!

Most of the ship’s time was spent surveying the coasts of South America and nearby islands, but it would also visit various Pacific islands, New Zealand, and Australia.  It was the Galapagos Islands that most impressed him.  There he found that finches had evolved a variety of beaks -- each suited to a particular food source.  Natural variation had somehow been selected to fit the ecological niches available on the tiny islands.

Upon returning, Darwin wrote several books based on his surveys on geology and the plant and animal species he had observed and collected.  He also published his journal as Journal of a Naturalist.  He notes that he was most impressed by the ways similar animals adapted to different ecologies.

From early on, Darwin recognized that selection was the principle men used so successfully when breeding animals.  What he needed now was an idea as to how nature could perform that task without the benefit of intelligence!

In 1838, he read a book by Malthus called Population.  Malthus introduced the idea that competition over limited resources would, in nature, keep populations stable.  He also warned that human populations, when straining resources, would suffer as well!

On January 29, 1839, Darwin married Emma Wedgewood, a cousin.  They lived in London for a few years, then settled in the village of Down, 15 miles outside London, where they lived the rest of their lives.  Darwin began suffering from an illness he had probably contracted from an insect bite in the Andes many years before.  Darwin’s son, Francis, later could not say enough about his mother’s dedication to his father’s well-being.  Without her, he would have been considerably less productive.  They would go on to have two daughters and five sons.

Darwin wrote a sketch of his theory in 1842.  In 1844, he wrote in a letter “At last gleams of light have come, and I am almost convinced (quite contrary to the opinion I started with) that species are not (it is like confessing a murder) immutable.”

He was about half done with a full exposition of his ideas when he received an essay from A. R. Wallace, with a request for comments.  The essay outlined a theory of natural selection!  Wallace, too, had read Malthus, and in 1858, while sick from fever, had the whole idea come to him in one flash.  Darwin, in his reluctance, had postponed revealing his ideas to the scientific public for 20 years!

Darwin forwarded the essay to his friend, Sir Charles Lyell of the Geological Society, as Wallace had requested.  Lyell sent the essay on, with an essay by Darwin, for presentation at a scientific conference.

The point they jointly made was clear:  Just like men can exaggerate one or another minor variation by selectively breeding dogs or cattle, so nature selects similar variations -- by only permitting the most successful variations to survive and reproduce in the struggle over limited resources.  Although the changes would be slight and slow, the millennia would permit the obvious diversity of nature!  Darwin named this natural selection.

In 1859, Darwin finally published his master work, on the Origin of Species by Means of Natural Selection.  The book was an instant success.  There was also, of course, a great deal of debate -- mostly concerning the contrast with traditional religious explanations of the natural world.

Natural selection was often confused with an earlier idea of the French naturalist Lamarck.  He suggested that characteristics acquired during an animal’s life were passed on to its offspring.  The famous example is how the constant stretching of the neck over many generations explains the giraffe’s unlikely physique.  This theory -- Lamarckianism -- was natural selection’s major competitor for decades to come!

In 1868, he published The Variation of Animals and Plants under Domestication.  Then, in 1871, he came out with The Descent of Man, and Selection in Relation to Sex.  This was really two books in one.  The second part is about sexual selection.  This is what accounts for, for example, the bright colors of many male birds:  Both the males' coloring and the attraction to the coloring on the part of the females during courtship are selected for because these variations benefit the offspring.

The Descent of Man portion of the book is a brief introduction to the idea that we, too, are the results of natural selection.  This part would lead to a lot of heated arguments!

In 1872, The Expression of Emotion was published.  This time, Darwin talks about the evolution of the signals that animals use to communicate -- and relates those signals to human emotional expression.  This is the first step towards what we now call sociobiology (and evolutionary psychology).

In addition to these influential books, Darwin also enjoyed studying and writing about plants.  In 1862, he wrote Fertilization of Orchids.  In 1875, he came out with both Climbing Plants and Insectivorous Plants.  In 1877, came Different Forms of Flowers on Plants of the Same Species.  In 1880, he wrote, with his son Francis, The Power of Movement in Plants.  And in 1881, he published the famous Formation of Vegetable Mould through the Action of Worms!

Charles Darwin died April 19, 1882.  He was buried in Westminster Abbey.  He was apparently a kind and gentle man, beloved by his family and friends alike.  Outside of his voyage on the Beagle, he rarely left his home in Down.  Reluctantly, he surrendered his religious beliefs and settled into an agnosticism that did not prevent him from participating with his parish in charitable works.